Lecture—29 September 2003

 

Topic—Metamerism

 

Some definitions-

Benthic environment—the environment at the bottom of a lake or ocean.

Some subdivisions

Epibenthic or epifaunal—creatures that live on top of the substrate

Infaunal—creatures that live in the substrate.

 

Before the evolution of metameric creatures, most organisms lived on top of the substrate.  After the invention of metamerism, creatures were able to move into the substrate.  What enabled them to do so was the fact that no longer did the organism have to contract it whole body for movement.  Individual segments could be contracted and expanded.  This allowed for regional patterns of body movement. Typicaly in an animal such as an earth worm a wave of contraction passes down the body.  In as much as each segment has bristles that anchor that segment, this —peristalsis

 

Nematodes, acanthocephalans, arthropods

Chitin in the outer body wall—complex mucopolysacaride

 

For growth to occur, the outer layer of chitin has to be shed—ecdysis

 

Homeobox genes, rRNA

 

Acoels—they’re not platyhelminthes

 

Whether things shed or don’t shed

 

Ecdysozoa—arthropods, aschelminthes, etc.

 

Changes in environments

 

Cambrian—organisms that live in aqueous environments

 

Movement into freshwater

 

Ionic balance and osmotic balance

 

Movement on to land