My major interests are:

  1. Marine invertebrate ecology
  2. Geographic information systems, remote sensing and landscape ecology
  3. Evolutionary biology
  4. Ecology of the northern temperate forest
  5. Biology of cephalopods
  6. History of American science, particularly the establishment and growth of biological field stations
  7. History of oceanography
  8. Human use of wilderness
  9. Use of computers and multimedia in teaching